The Shadow Leech Hive

The Shadow LeechShadow LeechMutated Kraata designed to drain light and corruption. Created by Mutran and allies, they transformed dozens of Matoran before the Energy Storm wiped them out. Hive was a bioweapons facility constructed in Karda NuiKarda NuiThe vast inner cavern at the core of the Matoran Universe, original home of the Av-Matoran, site of the Codrex and the final battle that decided the fate of Mata Nui. by the Brotherhood of MakutaMakuta (Species)Shadowy Antidermis beings and the Brotherhood's elite, Makuta commanded darkness and shaped the Matoran Universe's fate.. Designed and operated by MakutaMakuta (Species)Shadowy Antidermis beings and the Brotherhood's elite, Makuta commanded darkness and shaped the Matoran Universe's fate. MutranMutranMutran is a Makuta of the Brotherhood of Makuta who serves as the Makuta of Voya Nui and is among the most prolific Rahi creators within the organization. Hailing from the Southern Islands, he wears the Great Shelek and commands the powers of Shadow and Silence alongside the full range of Kraata abilities, wielding a Shadow Blade, Shadow Spear, and Tridax Pod in combat. with VicanVicanVican is a mutated Le-Matoran who previously served as a Shadow Matoran and as an assistant to Mutran. He wears a powerless Shelek, wields claws, and bore the black and lime coloring of his time as a Shadow Matoran. Connected to the element of Air, Vican resides on Spherus Magna. as forced labor, the hive served as the breeding centre for the Shadow Leeches deployed against the Av-MatoranAv-MatoranThe Light-element Matoran, prototypes of their species, who colonised Karda Nui from the Southern Continent and built the Stalactite Villages above the Swamp of Secrets. during the Karda NuiKarda NuiThe vast inner cavern at the core of the Matoran Universe, original home of the Av-Matoran, site of the Codrex and the final battle that decided the fate of Mata Nui. campaign.

Origin#

The Shadow LeechShadow LeechMutated Kraata designed to drain light and corruption. Created by Mutran and allies, they transformed dozens of Matoran before the Energy Storm wiped them out. Hive was established in the flooded core of Karda NuiKarda NuiThe vast inner cavern at the core of the Matoran Universe, original home of the Av-Matoran, site of the Codrex and the final battle that decided the fate of Mata Nui., the region rendered into the Swamp of SecretsSwamp of SecretsThe Swamp of Secrets was the mutagen-saturated wetland that filled the floor of Karda Nui after the Great Cataclysm, serving as a Brotherhood stronghold until the Energy Storms. after the Great Cataclysm. When MutranMutranMutran is a Makuta of the Brotherhood of Makuta who serves as the Makuta of Voya Nui and is among the most prolific Rahi creators within the organization. Hailing from the Southern Islands, he wears the Great Shelek and commands the powers of Shadow and Silence alongside the full range of Kraata abilities, wielding a Shadow Blade, Shadow Spear, and Tridax Pod in combat. developed the Shadow LeechShadow LeechMutated Kraata designed to drain light and corruption. Created by Mutran and allies, they transformed dozens of Matoran before the Energy Storm wiped them out., a mutated kraata capable of draining light from living beings, the Brotherhood required a production site. The swamp, contaminated with Pit mutagen and concealed beneath the dome, offered the necessary isolation. Three MakutaMakuta (Species)Shadowy Antidermis beings and the Brotherhood's elite, Makuta commanded darkness and shaped the Matoran Universe's fate. descended into its waters, were mutated into their insectoid MistikaMistikaMistika was the designation given to the Toa Nuva and Makuta who fought in the Swamp of Secrets at the floor of Karda Nui during the final campaign. forms, and established the base that became the hive.

MutranMutranMutran is a Makuta of the Brotherhood of Makuta who serves as the Makuta of Voya Nui and is among the most prolific Rahi creators within the organization. Hailing from the Southern Islands, he wears the Great Shelek and commands the powers of Shadow and Silence alongside the full range of Kraata abilities, wielding a Shadow Blade, Shadow Spear, and Tridax Pod in combat. was the originator of the Shadow LeechShadow LeechMutated Kraata designed to drain light and corruption. Created by Mutran and allies, they transformed dozens of Matoran before the Energy Storm wiped them out. project. He referred to the creatures as among his "fabulous creatures" and tested them on a MatoranMatoranThe most abundant species in the Matoran Universe, chosen by Mata Nui to inhabit and maintain his cosmic body. in his territory before scaling up production. His assistant VicanVicanVican is a mutated Le-Matoran who previously served as a Shadow Matoran and as an assistant to Mutran. He wears a powerless Shelek, wields claws, and bore the black and lime coloring of his time as a Shadow Matoran. Connected to the element of Air, Vican resides on Spherus Magna. maintained the breeding vats, but the operation was directed by MutranMutranMutran is a Makuta of the Brotherhood of Makuta who serves as the Makuta of Voya Nui and is among the most prolific Rahi creators within the organization. Hailing from the Southern Islands, he wears the Great Shelek and commands the powers of Shadow and Silence alongside the full range of Kraata abilities, wielding a Shadow Blade, Shadow Spear, and Tridax Pod in combat. himself. The hive produced swarms of the leeches for deployment against the Av-MatoranAv-MatoranThe Light-element Matoran, prototypes of their species, who colonised Karda Nui from the Southern Continent and built the Stalactite Villages above the Swamp of Secrets. and other targets in Karda NuiKarda NuiThe vast inner cavern at the core of the Matoran Universe, original home of the Av-Matoran, site of the Codrex and the final battle that decided the fate of Mata Nui..

The hive's location took advantage of the swamp's mutagenic waters, the natural hazards surrounding it, and additional defenses including the sonic-screaming klakk. The position represented a shift in the Brotherhood's methods toward direct biological warfare and the engineering of corrupted populations.

The hive was suspended from the cavern ceiling by a living stone cord. The structure's eventual destruction came after Kirop, an Av-MatoranAv-MatoranThe Light-element Matoran, prototypes of their species, who colonised Karda Nui from the Southern Continent and built the Stalactite Villages above the Swamp of Secrets. captive, escaped and led the Toa NuvaToa Mata and Toa NuvaSix biomechanical heroes created by the Great Beings to awaken Mata Nui and defeat the Brotherhood, transformed once and tested across an entire age. to the site, after which Toa Ignika severed the cord and dropped the facility into the swamp.

Breeding operations#

MutranMutranMutran is a Makuta of the Brotherhood of Makuta who serves as the Makuta of Voya Nui and is among the most prolific Rahi creators within the organization. Hailing from the Southern Islands, he wears the Great Shelek and commands the powers of Shadow and Silence alongside the full range of Kraata abilities, wielding a Shadow Blade, Shadow Spear, and Tridax Pod in combat.'s laboratory occupied the heart of the hive. Bubbling vats containing developing Shadow LeechesShadow LeechMutated Kraata designed to drain light and corruption. Created by Mutran and allies, they transformed dozens of Matoran before the Energy Storm wiped them out. were its central feature. VicanVicanVican is a mutated Le-Matoran who previously served as a Shadow Matoran and as an assistant to Mutran. He wears a powerless Shelek, wields claws, and bore the black and lime coloring of his time as a Shadow Matoran. Connected to the element of Air, Vican resides on Spherus Magna., the MatoranMatoranThe most abundant species in the Matoran Universe, chosen by Mata Nui to inhabit and maintain his cosmic body. pressed into service, did not labor willingly; MutranMutranMutran is a Makuta of the Brotherhood of Makuta who serves as the Makuta of Voya Nui and is among the most prolific Rahi creators within the organization. Hailing from the Southern Islands, he wears the Great Shelek and commands the powers of Shadow and Silence alongside the full range of Kraata abilities, wielding a Shadow Blade, Shadow Spear, and Tridax Pod in combat. had taken him from a village under the pretense of offering adventure and assigned him to the cleanup of failed experiments and the maintenance of containment systems. MutranMutranMutran is a Makuta of the Brotherhood of Makuta who serves as the Makuta of Voya Nui and is among the most prolific Rahi creators within the organization. Hailing from the Southern Islands, he wears the Great Shelek and commands the powers of Shadow and Silence alongside the full range of Kraata abilities, wielding a Shadow Blade, Shadow Spear, and Tridax Pod in combat. characterized this arrangement as a fair exchange for the loss of VicanVicanVican is a mutated Le-Matoran who previously served as a Shadow Matoran and as an assistant to Mutran. He wears a powerless Shelek, wields claws, and bore the black and lime coloring of his time as a Shadow Matoran. Connected to the element of Air, Vican resides on Spherus Magna.'s light.

The Shadow LeechShadow LeechMutated Kraata designed to drain light and corruption. Created by Mutran and allies, they transformed dozens of Matoran before the Energy Storm wiped them out. itself was a kraata modified by MutranMutranMutran is a Makuta of the Brotherhood of Makuta who serves as the Makuta of Voya Nui and is among the most prolific Rahi creators within the organization. Hailing from the Southern Islands, he wears the Great Shelek and commands the powers of Shadow and Silence alongside the full range of Kraata abilities, wielding a Shadow Blade, Shadow Spear, and Tridax Pod in combat. to drain the light from living beings. The drain operated on both physical and moral planes, stripping away the inner light tied to clarity of judgment along with raw energy. TakanuvaTakanuvaHidden among the Ta-Matoran as Takua, the disguised Av-Matoran who became Takanuva was the sole Toa of Light produced in the Matoran Universe's full history. was rendered unconscious after a leech attached to him, weakened and left collapsed in the Archives. MutranMutranMutran is a Makuta of the Brotherhood of Makuta who serves as the Makuta of Voya Nui and is among the most prolific Rahi creators within the organization. Hailing from the Southern Islands, he wears the Great Shelek and commands the powers of Shadow and Silence alongside the full range of Kraata abilities, wielding a Shadow Blade, Shadow Spear, and Tridax Pod in combat. described his creations as spreading darkness and corruption. The breeding vats served as production lines for parasites engineered to destabilize forces of Light before direct engagement.

Security around the hive was substantial but not impenetrable. KopakaKopakaKopaka was the Toa Nuva of Ice, deputy commander of the Toa Mata, and one of the six warriors crafted on Artakha to defend the Matoran Universe., SolekSolekSolek was an Av-Matoran of Karda Nui who fought alongside the Toa Nuva against the Brotherhood of Makuta during the final battle for Mata Nui's awakening., and others breached it after Kirop led them inside. MutranMutranMutran is a Makuta of the Brotherhood of Makuta who serves as the Makuta of Voya Nui and is among the most prolific Rahi creators within the organization. Hailing from the Southern Islands, he wears the Great Shelek and commands the powers of Shadow and Silence alongside the full range of Kraata abilities, wielding a Shadow Blade, Shadow Spear, and Tridax Pod in combat. defended his position using illusions and mental assaults to turn the Toa against one another. The physical infrastructure held vats and binding systems strong enough to restrain prisoners until Toa Ignika passed through them and freed the captives. The Toa NuvaToa Mata and Toa NuvaSix biomechanical heroes created by the Great Beings to awaken Mata Nui and defeat the Brotherhood, transformed once and tested across an entire age. then destroyed the vats in a coordinated strike. The hive itself fell only after Ignika killed the living stone cord supporting it.

The Toa Nuva raid#

Kirop's escape from captivity provided the route to the hive. He had overheard MutranMutranMutran is a Makuta of the Brotherhood of Makuta who serves as the Makuta of Voya Nui and is among the most prolific Rahi creators within the organization. Hailing from the Southern Islands, he wears the Great Shelek and commands the powers of Shadow and Silence alongside the full range of Kraata abilities, wielding a Shadow Blade, Shadow Spear, and Tridax Pod in combat. discussing the Shadow LeechShadow LeechMutated Kraata designed to drain light and corruption. Created by Mutran and allies, they transformed dozens of Matoran before the Energy Storm wiped them out. vats and slipped out, unaware that he was being followed by KopakaKopakaKopaka was the Toa Nuva of Ice, deputy commander of the Toa Mata, and one of the six warriors crafted on Artakha to defend the Matoran Universe., SolekSolekSolek was an Av-Matoran of Karda Nui who fought alongside the Toa Nuva against the Brotherhood of Makuta during the final battle for Mata Nui's awakening., Tanma, PhotokPhotokPhotok was an Av-Matoran of Karda Nui who fought alongside Tanma and Solek in the resistance against the Brotherhood of Makuta during the final battle for Mata Nui's awakening., and the others. His flight led the Toa NuvaToa Mata and Toa NuvaSix biomechanical heroes created by the Great Beings to awaken Mata Nui and defeat the Brotherhood, transformed once and tested across an entire age. directly to the facility.

The Toa NuvaToa Mata and Toa NuvaSix biomechanical heroes created by the Great Beings to awaken Mata Nui and defeat the Brotherhood, transformed once and tested across an entire age. entered fast. While MutranMutranMutran is a Makuta of the Brotherhood of Makuta who serves as the Makuta of Voya Nui and is among the most prolific Rahi creators within the organization. Hailing from the Southern Islands, he wears the Great Shelek and commands the powers of Shadow and Silence alongside the full range of Kraata abilities, wielding a Shadow Blade, Shadow Spear, and Tridax Pod in combat. engaged KopakaKopakaKopaka was the Toa Nuva of Ice, deputy commander of the Toa Mata, and one of the six warriors crafted on Artakha to defend the Matoran Universe. and tricked the others into attacking Toa Ignika, the remainder moved on the vats. When the other three MakutaMakuta (Species)Shadowy Antidermis beings and the Brotherhood's elite, Makuta commanded darkness and shaped the Matoran Universe's fate. arrived and took the group captive, PohatuPohatuPohatu was the Toa Mata and Toa Nuva of Stone, guardian of Po-Wahi, whose speed and steady humour anchored the team through every era of the Matoran Universe. weakened the floor beneath the MakutaMakuta (Species)Shadowy Antidermis beings and the Brotherhood's elite, Makuta commanded darkness and shaped the Matoran Universe's fate., activated his Mask of Speed to phase through his bonds, and freed the others. He then used the Kakama Nuva to destroy the hive. The strike removed the immediate stockpile of Shadow LeechesShadow LeechMutated Kraata designed to drain light and corruption. Created by Mutran and allies, they transformed dozens of Matoran before the Energy Storm wiped them out. available to the Brotherhood.

The strategic effect was significant though not complete. The raid disabled the active production line, but the hive itself was brought down only when Toa Ignika killed the living stone cord supporting it. Once severed, the structure dropped into the Swamp of SecretsSwamp of SecretsThe Swamp of Secrets was the mutagen-saturated wetland that filled the floor of Karda Nui after the Great Cataclysm, serving as a Brotherhood stronghold until the Energy Storms. below. The Energy StormEnergy Storm: The Raw-Energy Maelstroms of Karda NuiThe Energy Storms were vast maelstroms of raw energy that tore through Karda Nui whenever the Great Spirit Mata Nui woke, devastating the region while feeding power into his body. that followed Mata NuiMata Nui (Location)An island created by accident on the Great Spirit's face, home to the Matoran for a thousand years.'s awakening erased whatever remained.

Collapse#

The hive's structural collapse began with Toa Ignika's destruction of the living stone cord that suspended the facility from the cavern roof. The cord was a semi-sentient organism, tied to the wider mutagenic ecosystem of Karda NuiKarda NuiThe vast inner cavern at the core of the Matoran Universe, original home of the Av-Matoran, site of the Codrex and the final battle that decided the fate of Mata Nui.. When Ignika severed it, the hive broke loose and dropped into the Swamp of SecretsSwamp of SecretsThe Swamp of Secrets was the mutagen-saturated wetland that filled the floor of Karda Nui after the Great Cataclysm, serving as a Brotherhood stronghold until the Energy Storms..

The impact rendered the facility inoperable. Vats holding developing Shadow LeechesShadow LeechMutated Kraata designed to drain light and corruption. Created by Mutran and allies, they transformed dozens of Matoran before the Energy Storm wiped them out. ruptured on impact, ending MutranMutranMutran is a Makuta of the Brotherhood of Makuta who serves as the Makuta of Voya Nui and is among the most prolific Rahi creators within the organization. Hailing from the Southern Islands, he wears the Great Shelek and commands the powers of Shadow and Silence alongside the full range of Kraata abilities, wielding a Shadow Blade, Shadow Spear, and Tridax Pod in combat.'s operations there. The Toa NuvaToa Mata and Toa NuvaSix biomechanical heroes created by the Great Beings to awaken Mata Nui and defeat the Brotherhood, transformed once and tested across an entire age., guided to the hive by Kirop, had already destroyed multiple breeding chambers. With the structure broken and submerged in a biologically hostile environment, any surviving leeches would have been exposed to swamp mutagen, likely corrupting or incapacitating them. The Brotherhood lost a critical node in their Karda NuiKarda NuiThe vast inner cavern at the core of the Matoran Universe, original home of the Av-Matoran, site of the Codrex and the final battle that decided the fate of Mata Nui. campaign and did not rebuild it.

Final obliteration came with the Energy StormEnergy Storm: The Raw-Energy Maelstroms of Karda NuiThe Energy Storms were vast maelstroms of raw energy that tore through Karda Nui whenever the Great Spirit Mata Nui woke, devastating the region while feeding power into his body.. TakanuvaTakanuvaHidden among the Ta-Matoran as Takua, the disguised Av-Matoran who became Takanuva was the sole Toa of Light produced in the Matoran Universe's full history. had warned the Toa NuvaToa Mata and Toa NuvaSix biomechanical heroes created by the Great Beings to awaken Mata Nui and defeat the Brotherhood, transformed once and tested across an entire age. that Mata NuiMata Nui (Location)An island created by accident on the Great Spirit's face, home to the Matoran for a thousand years.'s awakening would unleash a violent surge of energy through Karda NuiKarda NuiThe vast inner cavern at the core of the Matoran Universe, original home of the Av-Matoran, site of the Codrex and the final battle that decided the fate of Mata Nui.. The storm consumed everything suspended in the dome, including the remains of the hive, surviving leeches, and the MakutaMakuta (Species)Shadowy Antidermis beings and the Brotherhood's elite, Makuta commanded darkness and shaped the Matoran Universe's fate. outposts in the region. The hive's wreckage, already weakened and submerged, was reduced to inert matter.

Strategic role#

The hive occupied a central place in the Brotherhood's plan to dominate Karda NuiKarda NuiThe vast inner cavern at the core of the Matoran Universe, original home of the Av-Matoran, site of the Codrex and the final battle that decided the fate of Mata Nui.. The three MistikaMistikaMistika was the designation given to the Toa Nuva and Makuta who fought in the Swamp of Secrets at the floor of Karda Nui during the final campaign.-form MakutaMakuta (Species)Shadowy Antidermis beings and the Brotherhood's elite, Makuta commanded darkness and shaped the Matoran Universe's fate. who landed in the flooded core established the camp that became the breeding and deployment centre for the leeches. The leeches were intended to possess and convert the Av-MatoranAv-MatoranThe Light-element Matoran, prototypes of their species, who colonised Karda Nui from the Southern Continent and built the Stalactite Villages above the Swamp of Secrets. and ultimately the Toa, turning the inhabitants of Karda NuiKarda NuiThe vast inner cavern at the core of the Matoran Universe, original home of the Av-Matoran, site of the Codrex and the final battle that decided the fate of Mata Nui. into agents of darkness.

The destruction of the hive followed swiftly on its rise to operational status. Kirop's escape, held in a hut near the hive, exposed the MakutaMakuta (Species)Shadowy Antidermis beings and the Brotherhood's elite, Makuta commanded darkness and shaped the Matoran Universe's fate.'s plans. He led the Toa NuvaToa Mata and Toa NuvaSix biomechanical heroes created by the Great Beings to awaken Mata Nui and defeat the Brotherhood, transformed once and tested across an entire age., Tanma, SolekSolekSolek was an Av-Matoran of Karda Nui who fought alongside the Toa Nuva against the Brotherhood of Makuta during the final battle for Mata Nui's awakening., PhotokPhotokPhotok was an Av-Matoran of Karda Nui who fought alongside Tanma and Solek in the resistance against the Brotherhood of Makuta during the final battle for Mata Nui's awakening., and eventually Toa Ignika back to the facility. The raid struck the Brotherhood's operational core. The Toa NuvaToa Mata and Toa NuvaSix biomechanical heroes created by the Great Beings to awaken Mata Nui and defeat the Brotherhood, transformed once and tested across an entire age. destroyed vats and disrupted the production line; Toa Ignika then severed the cord, dropping the structure into the swamp. The Energy Storm finished the destruction.

The hive's fall marked a wider failure for the Brotherhood. The MakutaMakuta (Species)Shadowy Antidermis beings and the Brotherhood's elite, Makuta commanded darkness and shaped the Matoran Universe's fate. had counted on using the swamp's mutagenic waters to their advantage, despite being themselves locked into MistikaMistikaMistika was the designation given to the Toa Nuva and Makuta who fought in the Swamp of Secrets at the floor of Karda Nui during the final campaign. forms by the same mutagen. The same conditions that had reshaped them also rendered their operations vulnerable; once the Toa NuvaToa Mata and Toa NuvaSix biomechanical heroes created by the Great Beings to awaken Mata Nui and defeat the Brotherhood, transformed once and tested across an entire age. breached the facility, the Brotherhood's hold on Karda NuiKarda NuiThe vast inner cavern at the core of the Matoran Universe, original home of the Av-Matoran, site of the Codrex and the final battle that decided the fate of Mata Nui. began to break apart.

Source contradictions#

The destruction of the hive appears in two distinct accounts. The first, from BIONICLEBIONICLE: Matoran AdventuresA 2002 GBA platformer where you play as one of six Matoran fighting Bohrok across Mata Nui's islands. Legends 9: Shadows in the Sky, describes the Toa NuvaToa Mata and Toa NuvaSix biomechanical heroes created by the Great Beings to awaken Mata Nui and defeat the Brotherhood, transformed once and tested across an entire age.'s raid and Toa Ignika's severing of the stone cord, sending the hive crashing into the Swamp of SecretsSwamp of SecretsThe Swamp of Secrets was the mutagen-saturated wetland that filled the floor of Karda Nui after the Great Cataclysm, serving as a Brotherhood stronghold until the Energy Storms.. The second, recorded in the Swamp of SecretsSwamp of SecretsThe Swamp of Secrets was the mutagen-saturated wetland that filled the floor of Karda Nui after the Great Cataclysm, serving as a Brotherhood stronghold until the Energy Storms. serial, states that the hive was destroyed by the Energy Storm at Mata NuiMata Nui (Location)An island created by accident on the Great Spirit's face, home to the Matoran for a thousand years.'s awakening. Both accounts are canonical, and neither was retracted. The most consistent reading is that the fall into the swamp rendered the hive inoperable while the Energy Storm provided the final material destruction.

The role of Toa Ignika versus the Energy Storm also differs between accounts. The Legends version assigns direct responsibility to Ignika for the fall, while BIONICLEBIONICLE: Matoran AdventuresA 2002 GBA platformer where you play as one of six Matoran fighting Bohrok across Mata Nui's islands. Ignition 14: Endgame frames the Energy Storm as the inevitable outcome of Mata NuiMata NuiBuilt by the Great Beings to restore Spherus Magna, Mata Nui sustained an entire civilisation inside his body, was deposed by Teridax, and reclaimed his purpose as a mortal on Bara Magna.'s awakening, capable of erasing all structures in Karda NuiKarda NuiThe vast inner cavern at the core of the Matoran Universe, original home of the Av-Matoran, site of the Codrex and the final battle that decided the fate of Mata Nui. regardless of prior damage. Ignika's intervention was therefore decisive but not exclusive; the storm would have destroyed the hive in any case.

The Swamp of SecretsSwamp of SecretsThe Swamp of Secrets was the mutagen-saturated wetland that filled the floor of Karda Nui after the Great Cataclysm, serving as a Brotherhood stronghold until the Energy Storms. serial confirms that the Energy Storm incinerated the swamp and killed all the MakutaMakuta (Species)Shadowy Antidermis beings and the Brotherhood's elite, Makuta commanded darkness and shaped the Matoran Universe's fate. caught there, who could not escape in time. Since the hive lay in the swamp by that point, the storm would have consumed it along with the MakutaMakuta (Species)Shadowy Antidermis beings and the Brotherhood's elite, Makuta commanded darkness and shaped the Matoran Universe's fate.. The sequence of damage is therefore cumulative: the Toa NuvaToa Mata and Toa NuvaSix biomechanical heroes created by the Great Beings to awaken Mata Nui and defeat the Brotherhood, transformed once and tested across an entire age. disabled the production line, Ignika dropped the structure, and the storm reduced it to inert matter.

Significance#

The Shadow LeechShadow LeechMutated Kraata designed to drain light and corruption. Created by Mutran and allies, they transformed dozens of Matoran before the Energy Storm wiped them out. Hive represented a shift in the Brotherhood of MakutaMakuta (Species)Shadowy Antidermis beings and the Brotherhood's elite, Makuta commanded darkness and shaped the Matoran Universe's fate.'s methods toward industrial biological warfare. MutranMutranMutran is a Makuta of the Brotherhood of Makuta who serves as the Makuta of Voya Nui and is among the most prolific Rahi creators within the organization. Hailing from the Southern Islands, he wears the Great Shelek and commands the powers of Shadow and Silence alongside the full range of Kraata abilities, wielding a Shadow Blade, Shadow Spear, and Tridax Pod in combat.'s work on Shadow Leeches and earlier on VultrazVultrazVultraz is a Shadow Matoran, formerly a Ta-Matoran, who wears a powerless Mask of Scavenging and carries a sword. Dark red and black in coloration, he possesses an active power of Shadow while his former affinity for Fire remains inactive. He resides in a parallel universe. demonstrated the MakutaMakuta (Species)Shadowy Antidermis beings and the Brotherhood's elite, Makuta commanded darkness and shaped the Matoran Universe's fate.'s capacity to develop weapons capable of stripping light from living beings on a scalable basis. The hive made this capability operational.

The facility was destroyed before its full output could be deployed. TakanuvaTakanuvaHidden among the Ta-Matoran as Takua, the disguised Av-Matoran who became Takanuva was the sole Toa of Light produced in the Matoran Universe's full history.'s incapacitation by a leech in the Archives demonstrated the weapon's effectiveness against even a Toa of LightToa of LightThe sole Toa wielding the Element of Light through the Avohkii mask, singular and feared by Makuta itself., showing that light could be drained passively without direct combat. The hive's destruction in the Karda NuiKarda NuiThe vast inner cavern at the core of the Matoran Universe, original home of the Av-Matoran, site of the Codrex and the final battle that decided the fate of Mata Nui. campaign removed this capability from the Brotherhood's arsenal and contributed to the wider collapse of their position in the region.