Krana

Krana were small organic symbiotes engineered by the Great BeingsThe Great BeingsScholar-inventors of Spherus Magna whose creations shaped the Matoran Universe and whose absence defined the conflicts that followed. and later mass-produced by the BahragBahragCahdok and Gahdok were the twin queens of the Bohrok, sealed in a Protodermis cage by the Toa Mata and later released to finish their cleansing of Mata Nui. to function as the control units of the BohrokBohrokHive-minded mechanical creatures designed to reshape worlds, the Bohrok represent a threat the Toa Mata never expected. swarms. Each BohrokBohrokHive-minded mechanical creatures designed to reshape worlds, the Bohrok represent a threat the Toa Mata never expected. was a fully mechanical chassis. The krana was the living component inside the head. Without a krana, a BohrokBohrokHive-minded mechanical creatures designed to reshape worlds, the Bohrok represent a threat the Toa Mata never expected. was an inert shell. With one, it was a member of an organized colony directed toward a single goal.

Form and biology#

A krana was a soft organic creature with a mask-like face and an adhesive underside, smaller than a MatoranMatoranThe most abundant species in the Matoran Universe, chosen by Mata Nui to inhabit and maintain his cosmic body.. It attached to the front of a BohrokBohrokHive-minded mechanical creatures designed to reshape worlds, the Bohrok represent a threat the Toa Mata never expected. and operated the chassis through the interface the BahragBahragCahdok and Gahdok were the twin queens of the Bohrok, sealed in a Protodermis cage by the Toa Mata and later released to finish their cleansing of Mata Nui. had designed for it. When the Onu-MatoranOnu-MatoranEarth-wielding Matoran of the deep who adapted to darkness and built the Matoran Universe's defensive cavalry. MavrahMavrahA Metru Nui Archivist who fled his city rather than let Turaga Dume cull his pet sea Rahi, drowned in a feeding frenzy a thousand years before the Great Cataclysm, and was rediscovered alive aboard the Red Star. examined a captured BohrokBohrokHive-minded mechanical creatures designed to reshape worlds, the Bohrok represent a threat the Toa Mata never expected. during his early research in the Matoran UniverseMatoran UniverseA civilization of biomechanical beings living inside a colossal robot, destroyed when Teridax seized control., he confirmed that the chassis contained no organic components of its own. The krana was the sole living element of the system.

The krana was not optional equipment. It was the engine. A BohrokBohrokHive-minded mechanical creatures designed to reshape worlds, the Bohrok represent a threat the Toa Mata never expected. without its krana became an inert shell that could be repurposed, reprogrammed, or simply abandoned. This established the strategic principle of the Bohrok WarBohrok WarThe 2002 swarm that nearly destroyed Mata Nui when Teridax triggered the Bohrok's awakening ahead of schedule.: the Toa MataToa Mata and Toa NuvaSix biomechanical heroes created by the Great Beings to awaken Mata Nui and defeat the Brotherhood, transformed once and tested across an entire age. were never required to destroy a BohrokBohrokHive-minded mechanical creatures designed to reshape worlds, the Bohrok represent a threat the Toa Mata never expected. outright. They were required to extract the krana before the krana could relocate to another host.

Krana were classified as a species in the catalogues of the Matoran UniverseMatoran UniverseA civilization of biomechanical beings living inside a colossal robot, destroyed when Teridax seized control.. They were engineered. They executed their function. They had no recorded capacity for reflection, dissent, or independent purpose.

The eight standard breeds#

Eight krana breeds were produced, distinguished by two-letter suffix and specialized function. A BohrokBohrokHive-minded mechanical creatures designed to reshape worlds, the Bohrok represent a threat the Toa Mata never expected. became the functional equivalent of its krana for the duration of the bond.

The Xa was the command krana. The Swarm Commander. A BohrokBohrokHive-minded mechanical creatures designed to reshape worlds, the Bohrok represent a threat the Toa Mata never expected. wearing a Xa could formulate complex tactical plans for the swarm. Without a Xa, a swarm acted as a flat wave. With one, it conducted operations.

The Za was the telepathic relay. The Squad Leader. A Za-bonded BohrokBohrokHive-minded mechanical creatures designed to reshape worlds, the Bohrok represent a threat the Toa Mata never expected. coordinated the swarm across the entire span of Mata Nui IslandMata Nui (Location)An island created by accident on the Great Spirit's face, home to the Matoran for a thousand years. by telepathic link. The Za was also the breed responsible for the assimilation of LewaLewaToa of Air, member of the Toa Mata and Toa Nuva, whose career was marked by repeated possession, body theft, and aerial campaigns from Le-Wahi to Karda Nui. and Turaga MatauMatauA former Le-Metru test driver who became a Toa Metru, Matau wielded air, illusion, and shape-shifting before sacrificing his Toa power to wake the sleeping Matoran. during the fall of Le-KoroLe-KoroLe-Koro is a village situated within Le-Wahi, inhabited by the Le-Matoran and home to Turaga Matau and Toa Lewa. The settlement was once held under the control of Teridax..

The Vu provided short-range flight. A Vu-piloted BohrokBohrokHive-minded mechanical creatures designed to reshape worlds, the Bohrok represent a threat the Toa Mata never expected. could hop ahead of the swarm to clear sightlines or retreat from a threat. The Vu was airborne before it was recognized.

The Ca was the shielding breed. Called Clearance, the Ca generated a defensive shield for itself and the units immediately around it. The front rank of a swarm absorbed hits the rest did not have to.

The Yo was the tunneling breed. The Mole. A Yo-bonded BohrokBohrokHive-minded mechanical creatures designed to reshape worlds, the Bohrok represent a threat the Toa Mata never expected. could excavate through almost any surface on Mata Nui IslandMata Nui (Location)An island created by accident on the Great Spirit's face, home to the Matoran for a thousand years., rendering terrain effectively irrelevant to the swarm's movement.

The Ja was the radar breed. It detected obstacles and threats at distance and relayed the data through the swarm. In recorded operations, a Ja inside a TahnokTahnokFire-element Bohrok swarms from Mata Nui, controlled by krana and armed with Fire Shields. detected a ToaToa (2001)Biomechanical heroes bonded to elemental powers and masks. Not born, but chosen and transformed. ambush ahead, and the corresponding Za relayed the warning across the swarm before contact was made.

The Su was raw strength. The Worker. A Su-bonded BohrokBohrokHive-minded mechanical creatures designed to reshape worlds, the Bohrok represent a threat the Toa Mata never expected. was assigned to brute-force tasks beyond the strength of the other breeds.

The Bo was the infrared breed. The Sentinel. A Bo could see in darkness and was deployed for night operations and tunnel work, particularly in the depths of the Bohrok NestBohrok NestThe subterranean chamber where the Bohrok slumbered beneath Mata Nui until Makuta's machinations awakened them..

Eight breeds operating through the Xa-and-Za command relay gave the swarm the organizational coherence of a single agent distributed across thousands of bodies.

Bonding and assimilation#

A krana would attach to any face. The BohrokBohrokHive-minded mechanical creatures designed to reshape worlds, the Bohrok represent a threat the Toa Mata never expected. chassis was a designed host, but the krana would bond equally well to a MatoranMatoranThe most abundant species in the Matoran Universe, chosen by Mata Nui to inhabit and maintain his cosmic body. or a ToaToa (2001)Biomechanical heroes bonded to elemental powers and masks. Not born, but chosen and transformed.. On a non-BohrokBohrokHive-minded mechanical creatures designed to reshape worlds, the Bohrok represent a threat the Toa Mata never expected. host, the krana sustained the wearer in a manner comparable to a KanohiKanohiMasks that grant powers and sustain life, they're the most vital artifact in the Matoran Universe.. If the BahragBahragCahdok and Gahdok were the twin queens of the Bohrok, sealed in a Protodermis cage by the Toa Mata and later released to finish their cleansing of Mata Nui. were active, the krana also routed the wearer's consciousness into the swarm.

This occurred at Le-KoroLe-KoroLe-Koro is a village situated within Le-Wahi, inhabited by the Le-Matoran and home to Turaga Matau and Toa Lewa. The settlement was once held under the control of Teridax., where the population was largely converted in a single sustained engagement. Bohrok-MatauMatauA former Le-Metru test driver who became a Toa Metru, Matau wielded air, illusion, and shape-shifting before sacrificing his Toa power to wake the sleeping Matoran., the TuragaTuragaFormer Toa who fulfilled their destiny and transformed into leaders. Turaga wield Noble Kanohi masks and weakened elemental powers, guiding their communities. under krana control, forced a krana onto LewaLewaToa of Air, member of the Toa Mata and Toa Nuva, whose career was marked by repeated possession, body theft, and aerial campaigns from Le-Wahi to Karda Nui. himself and conscripted him into the hive. TamaruTamaruTamaru was a Le-Matoran Forest Navigator of Le-Koro, Kongu's Kolhii partner, and the Le-Koro representative in the Chronicler's Company at the Battle of Kini-Nui. and KonguKonguKongu commanded Le-Koro's Gukko Force before a strike from the Kanohi Ignika transformed him into a Toa of Air who later served among the Toa Mahri. escaped conversion. TahuTahu, Toa of FireTahu served as Toa Mata, Toa Nuva, and Toa Mistika of Fire, leading his team from the first landing on Mata Nui to the destruction of the Rahkshi army on Bara Magna. was also briefly controlled by a krana during the same period, a fact he did not disclose to the rest of the team until much later.

The link operated as a two-way channel. The krana was not a remote control but a relay. A bonded host was connected through the Za telepathy network to every other krana in the swarm. Wearers who recovered from the bond reported acquiring information about the swarm's operations that they could not have known otherwise. LewaLewaToa of Air, member of the Toa Mata and Toa Nuva, whose career was marked by repeated possession, body theft, and aerial campaigns from Le-Wahi to Karda Nui.'s account, given after OnuaOnuaOnua was the Toa Mata and Toa Nuva of Earth, a quiet, deliberate warrior whose patience and underground mastery anchored his team across the Matoran Universe. removed his krana within the Bohrok NestBohrok NestThe subterranean chamber where the Bohrok slumbered beneath Mata Nui until Makuta's machinations awakened them., described receiving knowledge from the network during the brief period of conversion.

The implication of universal compatibility was that the BohrokBohrokHive-minded mechanical creatures designed to reshape worlds, the Bohrok represent a threat the Toa Mata never expected. were not a conventional military force. They were a colonization instrument with an interchangeable interface. The krana could be deployed on a mechanical chassis to produce a swarm or on a MatoranMatoranThe most abundant species in the Matoran Universe, chosen by Mata Nui to inhabit and maintain his cosmic body.'s face to convert the population.

A threatened krana could detach from its BohrokBohrokHive-minded mechanical creatures designed to reshape worlds, the Bohrok represent a threat the Toa Mata never expected. and leap at a nearby target, leaving the chassis behind. Bohrok VaBohrok VaScouts and support units for the Bohrok swarms, the Bohrok Va operated independently where the hive-mind Bohrok could not., the small scout units that accompanied the swarms, carried replacement krana into the field to restore disabled hosts. In recorded engagements of the Bohrok-KalBohrok-KalSix elite Bohrok designed to free the imprisoned Bahrag, they nearly defeated the Toa Nuva before the Toa turned their own powers against them. period, a Va inserted a replacement krana-kal into a Pahrak-KalPahrak-KalThe slowest Bohrok-Kal conceals immense destructive plasma power that grows beyond control and consumes itself. under combat conditions after the original had been removed.

The Bohrok War#

The mission assigned to the Toa MataToa Mata and Toa NuvaSix biomechanical heroes created by the Great Beings to awaken Mata Nui and defeat the Brotherhood, transformed once and tested across an entire age. by Turaga VakamaVakamaA Ta-Metru maskmaker transformed into the Toa Metru of Fire, leader of his team through the Visorak war, and Turaga of Ta-Koro for a thousand years. after the first BohrokBohrokHive-minded mechanical creatures designed to reshape worlds, the Bohrok represent a threat the Toa Mata never expected. raid on Ta-KoroTa-KoroTa-Koro was a settlement located in Ta-Wahi, home to the Ta-Matoran, Tahu, and Vakama, and formerly connected to Teridax. The settlement was ultimately destroyed. was to collect one of each of the eight breeds from each of the six swarms. The total requirement was forty-eight krana. The collected krana would then be slotted into receptacles in the central chamber of the Bohrok NestBohrok NestThe subterranean chamber where the Bohrok slumbered beneath Mata Nui until Makuta's machinations awakened them. to release the Exo-ToaExo-ToaToa-sized powered armor originally produced by the Brotherhood of Makuta, worn briefly by the Toa Mata and later deployed as autonomous enforcers. suits and open the route to the BahragBahragCahdok and Gahdok were the twin queens of the Bohrok, sealed in a Protodermis cage by the Toa Mata and later released to finish their cleansing of Mata Nui..

The mission shape was unusual for a ToaToa (2001)Biomechanical heroes bonded to elemental powers and masks. Not born, but chosen and transformed. team. The objective was extraction rather than destruction. Each ToaToa (2001)Biomechanical heroes bonded to elemental powers and masks. Not born, but chosen and transformed. took responsibility for the swarm operating in their assigned region. GaliGali, Toa of WaterThe Toa Nuva of Water and conscience of the Toa Mata, whose attunement to her element anchored the team across every campaign from Mata Nui to Karda Nui. and KopakaKopakaKopaka was the Toa Nuva of Ice, deputy commander of the Toa Mata, and one of the six warriors crafted on Artakha to defend the Matoran Universe. conducted a joint water-trap operation against the TahnokTahnokFire-element Bohrok swarms from Mata Nui, controlled by krana and armed with Fire Shields., drowning a swarm and diving in to remove the krana from the disabled units. The operation succeeded in part but nearly cost Gali her life when the surviving TahnokTahnokFire-element Bohrok swarms from Mata Nui, controlled by krana and armed with Fire Shields. melted a canyon wall and dropped a stone slab on her position.

The Le-KoroLe-KoroLe-Koro is a village situated within Le-Wahi, inhabited by the Le-Matoran and home to Turaga Matau and Toa Lewa. The settlement was once held under the control of Teridax. operation was the most costly. By the time LewaLewaToa of Air, member of the Toa Mata and Toa Nuva, whose career was marked by repeated possession, body theft, and aerial campaigns from Le-Wahi to Karda Nui. returned, the village had been converted in its entirety. LewaLewaToa of Air, member of the Toa Mata and Toa Nuva, whose career was marked by repeated possession, body theft, and aerial campaigns from Le-Wahi to Karda Nui. was himself converted, used against his own allies, and only freed when OnuaOnuaOnua was the Toa Mata and Toa Nuva of Earth, a quiet, deliberate warrior whose patience and underground mastery anchored his team across the Matoran Universe. reached him in the Bohrok NestBohrok NestThe subterranean chamber where the Bohrok slumbered beneath Mata Nui until Makuta's machinations awakened them. and removed the krana from his face.

The endgame occurred inside the central chamber. Forty-eight krana were placed in the receptacles. The locks released. Six Exo-ToaExo-ToaToa-sized powered armor originally produced by the Brotherhood of Makuta, worn briefly by the Toa Mata and later deployed as autonomous enforcers. suits unsealed and were taken by the Toa MataToa Mata and Toa NuvaSix biomechanical heroes created by the Great Beings to awaken Mata Nui and defeat the Brotherhood, transformed once and tested across an entire age. for the assault on Cahdok and GahdokGahdokOne of the twin Bahrag queens, Gahdok commanded the Bohrok swarms alongside Cahdok until the Toa Mata sealed them both in solid protodermis.. The ToaToa (2001)Biomechanical heroes bonded to elemental powers and masks. Not born, but chosen and transformed. discarded the suits when the queens proved beyond their effective use and sealed the BahragBahragCahdok and Gahdok were the twin queens of the Bohrok, sealed in a Protodermis cage by the Toa Mata and later released to finish their cleansing of Mata Nui. in a Toa SealToa SealA cage of Protodermis created by six Toa combining their Elemental Powers, used to trap powerful beings. of ProtodermisProtodermisThe engineered substance building the Matoran Universe, protodermis exists in countless refined states from mundane metal to diamond-hard crystal to the destruction-or-transformation force of energized form. instead. With the queens contained, the swarms ceased operation. The MatoranMatoranThe most abundant species in the Matoran Universe, chosen by Mata Nui to inhabit and maintain his cosmic body. of Mata Nui IslandMata Nui (Location)An island created by accident on the Great Spirit's face, home to the Matoran for a thousand years. collected and sealed away every recovered krana.

Krana-Kal and the Bohrok-Kal incident#

The Bohrok-KalBohrok-KalSix elite Bohrok designed to free the imprisoned Bahrag, they nearly defeated the Toa Nuva before the Toa turned their own powers against them. were six elite BohrokBohrokHive-minded mechanical creatures designed to reshape worlds, the Bohrok represent a threat the Toa Mata never expected. who had not been dispatched by the BahragBahragCahdok and Gahdok were the twin queens of the Bohrok, sealed in a Protodermis cage by the Toa Mata and later released to finish their cleansing of Mata Nui. and activated independently with the objective of freeing their queens. The krana they carried were krana-kal, a variant mutated by exposure to a mutagenic substance.

The krana-kal carried greater onboard processing capacity than the standard krana. Where standard krana required the Za relay for coordination, each Kal operated as a fully autonomous agent. The Bohrok-KalBohrok-KalSix elite Bohrok designed to free the imprisoned Bahrag, they nearly defeated the Toa Nuva before the Toa turned their own powers against them. did not require a swarm structure.

The krana-kal were also capable of speech once removed from a host. During the operation against Tahnok-KalTahnok-KalThe electric-powered leader of the Bohrok-Kal squad, designed to awaken the Bahrag., PohatuPohatuPohatu was the Toa Mata and Toa Nuva of Stone, guardian of Po-Wahi, whose speed and steady humour anchored the team through every era of the Matoran Universe. bounced lightning off a polished surface to disable the Kal, and the Toa NuvaToa Mata and Toa NuvaSix biomechanical heroes created by the Great Beings to awaken Mata Nui and defeat the Brotherhood, transformed once and tested across an entire age. removed the krana-kal. The krana-kal reported the Kal's planned destination. A separate krana-kal recovered earlier through a BoxorBoxorBuilt by Nuparu from a salvaged Gahlok shell, the Boxor was the first Matoran-engineered combat machine and a key resistance asset against the Bohrok swarms. ambush had similarly briefed the TuragaTuragaFormer Toa who fulfilled their destiny and transformed into leaders. Turaga wield Noble Kanohi masks and weakened elemental powers, guiding their communities. on the Kal's mission.

In the final phase of the Kal operation, VakamaVakamaA Ta-Metru maskmaker transformed into the Toa Metru of Fire, leader of his team through the Visorak war, and Turaga of Ta-Koro for a thousand years. used the Kanohi VahiKanohi VahiThe Kanohi Vahi was the Legendary Mask of Time, forged by Toa Metru Vakama from the six fused Great Disks of Metru Nui during the city's evacuation. to slow time around the Bohrok-KalBohrok-KalSix elite Bohrok designed to free the imprisoned Bahrag, they nearly defeated the Toa Nuva before the Toa turned their own powers against them. immediately before they could employ the stolen Nuva symbols to free the BahragBahragCahdok and Gahdok were the twin queens of the Bohrok, sealed in a Protodermis cage by the Toa Mata and later released to finish their cleansing of Mata Nui.. With the queens about to be released, the krana-kal turned silver and projected an impenetrable force field. The Kal were stopped only by GaliGali, Toa of WaterThe Toa Nuva of Water and conscience of the Toa Mata, whose attunement to her element anchored the team across every campaign from Mata Nui to Karda Nui.'s plan to redirect each Kal's own power against itself. Gahlok-KalGahlok-KalThe magnetic-powered Bohrok-Kal whose own power became its undoing when victory was within reach. was crushed by Exo-ToaExo-ToaToa-sized powered armor originally produced by the Brotherhood of Makuta, worn briefly by the Toa Mata and later deployed as autonomous enforcers. drawn in by its magnetism. Lehvak-KalLehvak-KalThe vacuum-powered Bohrok-Kal who got blasted into space by its own power, Lehvak-Kal represents the chaotic energy of the Bohrok-Kal team. accelerated itself into space. Tahnok-KalTahnok-KalThe electric-powered leader of the Bohrok-Kal squad, designed to awaken the Bahrag. sealed itself inside its own lightning. Kohrak-KalKohrak-KalThe Bohrok-Kal of Sonics, disintegrated by the very element it wielded. disintegrated from its own sonic output. Some krana-kal survived the implosions and were collected into containment.

The silver, force-field-projecting state of the krana-kal at the moment of near-success indicated that the krana lineage was capable of further engineered escalation. The BahragBahragCahdok and Gahdok were the twin queens of the Bohrok, sealed in a Protodermis cage by the Toa Mata and later released to finish their cleansing of Mata Nui. had not completed development of the design.

Empty shells and parallel designs#

A BohrokBohrokHive-minded mechanical creatures designed to reshape worlds, the Bohrok represent a threat the Toa Mata never expected. without its krana was, per the encyclopedic record, an empty shell suitable for reprogramming or training. After the Bohrok WarBohrok WarThe 2002 swarm that nearly destroyed Mata Nui when Teridax triggered the Bohrok's awakening ahead of schedule., the MatoranMatoranThe most abundant species in the Matoran Universe, chosen by Mata Nui to inhabit and maintain his cosmic body. of Mata Nui IslandMata Nui (Location)An island created by accident on the Great Spirit's face, home to the Matoran for a thousand years. employed a workforce of disarmed BohrokBohrokHive-minded mechanical creatures designed to reshape worlds, the Bohrok represent a threat the Toa Mata never expected. in the reconstruction of the villages. The chassis were salvageable. The krana were not.

The FohrokFohrokManufactured Bohrok-analogues built by the Nynrah Ghosts for the Brotherhood of Makuta, destroyed by the Toa Hagah after rebelling against their handlers., so-called "fake BohrokBohrokHive-minded mechanical creatures designed to reshape worlds, the Bohrok represent a threat the Toa Mata never expected.," were a parallel design not controlled by krana. The Dark HuntersThe Dark HuntersA mercenary organization founded on Odina by The Shadowed One and Ancient, contracted across the Matoran Universe for assassination and theft, and ultimately reduced to a battered remnant by the fall of the universe itself. fielded them in their forces. They were functional but lacked the krana interface entirely, and operated through whatever substitute control system their engineers had improvised.

BoxorBoxorBuilt by Nuparu from a salvaged Gahlok shell, the Boxor was the first Matoran-engineered combat machine and a key resistance asset against the Bohrok swarms. units, the small mech suits constructed by Onu-MatoranOnu-MatoranEarth-wielding Matoran of the deep who adapted to darkness and built the Matoran Universe's defensive cavalry. during the Bohrok WarBohrok WarThe 2002 swarm that nearly destroyed Mata Nui when Teridax triggered the Bohrok's awakening ahead of schedule., gave the MatoranMatoranThe most abundant species in the Matoran Universe, chosen by Mata Nui to inhabit and maintain his cosmic body. a means of removing krana from disabled BohrokBohrokHive-minded mechanical creatures designed to reshape worlds, the Bohrok represent a threat the Toa Mata never expected. without exposing themselves to a krana jump. The same tool was used to ambush Bohrok-KalBohrok-KalSix elite Bohrok designed to free the imprisoned Bahrag, they nearly defeated the Toa Nuva before the Toa turned their own powers against them. in the later conflict.

Every krana sealed into MatoranMatoranThe most abundant species in the Matoran Universe, chosen by Mata Nui to inhabit and maintain his cosmic body. containment after the war remained a sleeping organism. The chassis were stored in the ArchivesThe Archives of Metru NuiThe Archives of Onu-Metru were the central repository of Matoran knowledge and Rahi specimens, maintained by the Archivist Corps beneath the city's surface.. The krana were stored separately. Both populations remained available for reactivation under the right conditions. The Order of Mata NuiThe Order of Mata NuiThe covert organization founded by Helryx to serve the Great Spirit directly, operating in concealment from a fortress on Daxia and waging the open war against the Brotherhood of Makuta that decided the fate of the universe., through AxonnAxonnAxonn served as an arbitrator under the Hand of Artakha and later as an Order of Mata Nui agent, guarding the Kanohi Ignika on Voya Nui for thousands of years., retained the option of releasing the BahragBahragCahdok and Gahdok were the twin queens of the Bohrok, sealed in a Protodermis cage by the Toa Mata and later released to finish their cleansing of Mata Nui. and reactivating the swarms as a strategic contingency. The contingency was never fully exercised, though BohrokBohrokHive-minded mechanical creatures designed to reshape worlds, the Bohrok represent a threat the Toa Mata never expected. released from the Metru NuiMetru NuiThe city beneath Mata Nui functions as the Great Spirit's brain, housing the Matoran whose labor keeps the god alive. ArchivesThe Archives of Metru NuiThe Archives of Onu-Metru were the central repository of Matoran knowledge and Rahi specimens, maintained by the Archivist Corps beneath the city's surface. during the Destiny WarThe Destiny WarThe open conflict between the Order of Mata Nui and the Brotherhood of Makuta that ended fifty thousand years of operational secrecy and broke the Brotherhood's last assault on Metru Nui. demonstrated that the activation chain remained functional.

Status#

The standing record describes krana as a controlled but not eliminated population. The BahragBahragCahdok and Gahdok were the twin queens of the Bohrok, sealed in a Protodermis cage by the Toa Mata and later released to finish their cleansing of Mata Nui. remained sealed for the duration of the Matoran UniverseMatoran UniverseA civilization of biomechanical beings living inside a colossal robot, destroyed when Teridax seized control. era. Without their queens, the swarms could not be coordinated, but the individual krana retained their function and could in principle be deployed against any compatible host. The chassis were inert. The relays were silent. The colony, in storage form, persisted.